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Lady Rose is the niece of Robert Crawley, Earl of Grantham. How she is related to him is currently unknown, but as Rosamund and Marmaduke had no children[3] it is most likely through Cora.

References

  1. The article, which can be seen here http://www.dailymail.co.uk/tvshowbiz/article-2156137/Fast-Girl-Lily-James-run-big-screen-straight-ITVs-hit-drama-Downton-Abbey.html?ito=feeds-newsxml states that most of her scenes are at the end of Series 3, and given that she looks about 20 and the end of Series 3 is set in 1921, we can give a rough estimate of her birthdate, which would be around 1901.
  2. In the article announcing that Lily James has joined Downton she states that she "likes being a brunette again" so Rose MUST be brown haired; the article can be seen here http://www.dailymail.co.uk/tvshowbiz/article-2156137/Fast-Girl-Lily-James-run-big-screen-straight-ITVs-hit-drama-Downton-Abbey.html?ito=feeds-newsxml.
  3. When Sybil states that she feels sorry for her aunt, Mary states that she doesn't and says that she'd love to "be alone in a big house in Eaton Squarw with all that money"'; this means that Rosamund is alone after being widowed and has no children, so Rose CANNOT be Rosamund's daughter otherwise Rosamund would NOT be alone, and thus, must be a niece of Cora and Robert would be her uncle by marriage
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